Institute of Science

Faculty of Science 



  • Ph.D., University of Virginia
  • M.S., Tokyo Metropolitan University
  • Bachelor of Science, Tokyo Metropolitan University

Field Of Study

  • Ecology and environment, Evolutionary ecology
  • Biodiversity and systematics, Phylogenetic systematics
  • Evolutionary biology, Evolutionary genetics


  • 2011
    Professor, Department of Biology, Shinshu University
  • 1999
    Associate Professor, Department of Biology, Shinshu University
  • 1995 - 1996
    Field Museum of Natural History (Chicago), Bishop Museum (Honolulu)
  • 1990
    Smithonian Institute
  • 1990
    Associate Professor, Tokyo Metropolitan College
  • 1989
    Research Associate, School of Environmental Sciences, University of Virginia/ Virginia Museum of Natural History

Educational Background

  • 1988, University of Virginia, School of Arts and Sciences, Biology
  • 1982 - 1983, Rotary Scholar, University of Virginia
  • 1978, Tokyo Metropolitan University, Faculty of Science, Biology
  • 1980, Tokyo Metropolitan University, School of Sciences, Biology
Research activity information


  • 2009
    ZoologicalSociety of Japan, Zoological Science Award


  • Left–right reversal recurrently evolved regardless of diaphanous-related formin gene duplication or loss in snails
    Takeshi Noda, Noriyuki Satoh, Edmund Gittenberger, Takahiro Asami
    Journal of Molecular Evolution, 2023, Refereed
  • Two subspecies of the snail mite Riccardoella (Proriccardoella) reaumuri Fain & van Goethem, 1986 (Acari, Prostigmata, Ereynetidae) from Japan
    Waki, T., Motochin, R., Asami, T., Shimano, S.
    Systematic & Applied Acarology, 27, 839-854, 2022, Refereed電子ジャーナル
  • Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue
    Páll-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T., Ablett, J. D.
    Zookeys, 981, 1-220, 2020, Refereed
  • Riccardoella mites infecting six land snail species, referred to in the Red Data List and the Red Data Book
    Waki T, Shimano S, Asami T
    Taxa,, 45, 34-39, 2019, Refereed
  • Subspecies description rates are higher in morphologically complex land snails
    Páll-Gergely B, Asami T, Sólymos P
    Zoologica Scripta, 48, 185-193, 2019, Refereed
  • Evolution of a dextral lineage by left-right reversal in Cristataria (Gastropoda, Pulmonata, Clausiliidae)
    Páll-Gergely P, Szekeres M, Fehér Z, Asami T, Harl J
    Journal of Zoological Systematics and Evolutionary Research, 57, 520-526, 2019, Refereed
  • Three phyla—Two type specimens—One shell: History of a snail shell revealed by modern imaging technology
    Schwaha T, Ruthensteiner B., Melzer R R, Asami T, Páll‐Gergely B
    Journal of Zoological Systematics and Evolutionary Research, 57, 527-533, 2019, Refereed
  • Realtime Social Networking Service rapidly reveals distributions of non-indigenous land snails in a European capital
    Páll-Gergely B, Majoros G, Domokos T, Juhász A, Turóci Á, Badacsonyi L, Fekete J, Asami T
    BioInvasions Records, 8, 782–792, 2019, Refereed
  • Heterochirality results from reduction of maternal diaph expression in a terrestrial pulmonate snail
    Noda T, Satoh N, Asami T
    Zoology Letters, 5(, 2019, Refereed
  • Riccardoella reaumuri Fain and van Goethem, 1986 (Acariformes: Trombidiformes: Ereynetidae) from terrestrial mollusks in Okinawajima Island, as first record from Japan
    Waki T, Shimano S, Asami T, Miyai T, Sasaki T
    The biological magazine Okinawa, 56, 27-31, 2018, Refereed
  • Enantiomorphs and taxonomy of three conchological species in flat-shelled snails Trichocathaica (Pulmonata: Camaenidae)
    Páll-Gergely B, Hunyadi A, Asami T
    Zookeys, 810, 19–44, 2018, Refereed
  • Taxonomical notes on Chinese camaenids with description of three new species (Gastropoda: Pulmonata)
    Wu, M, Asami, T
    Molluscan Research, 38, 137-148,, 2018, Refereed電子ジャーナル
  • A new species of Camaenidae (Gastropoda, Pulmonata) from Nusa Penida and Lombok Islands, Indonesia: novelty in a well-known fauna
    Páll-Gergely, B, Otani, JU, Hosoda, T, Asami, T, Harl, J
    Molluscan Research, 38, 41-49,, 2018, Refereed電子ジャーナル
  • A new species of Metalycaeus hitherto undistinguished from M. vinctus (Pilsbry, 1902) (Gastropoda: Cyclophoroidea: Alycaeidae)
    Páll-Gergely, B, Asami, T
    Venus, 75, 1–16,, 2017, Refereed
  • Molecular systematics of the land snail family Orculidae reveal paraphyly and deep splits within the clade Orthurethra (Gastropoda: Pulmonata)
    Harl, J, Haring, E, Sittenthaler, M, Sattmann, H, Asami, T, Páll-Gergely, B
    Zoological Journal of the Linnean Society, 181, 778–794,, 2017, RefereedWebofScience電子ジャーナル
  • Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
    Páll-Gergely, B, Hunyadi, A, Đỗ, Đ S, Naggs, F, Asami, T
    Zootaxa,, 4331, 1–124, 2017, RefereedWebofScience電子ジャーナル
  • Breathing device of a new Streptaulus species from Vietnam extends understanding of the function and structure of respiratory tubes in cyclophoroids (Gastropoda: Caenogastropoda: Pupinidae)
    Páll-Gergely, B, Gargominy, O, Fontaine, B, Asami, T
    Journal of Molluscan Studies, 83, 243-248,, 2017, RefereedWebofScience電子ジャーナル
  • A new diplommatinid genus and two new species from the Philippines (Gastropoda, Caenogastropoda, Cyclophoroidea)
    Páll-Gergely, B, Hunyadi, A, Asami, T
    Zookeys, 678, 1-10,, 2017, RefereedWebofScience電子ジャーナル
  • Three new species and a new genus of Hypselostomatidae (Gastropoda: Pulmonata) from Cong Troi Cave, Northern Vietnam
    Páll-Gergely B, Jochum, A, Asami, T
    Acta Zoologica Academiae Scientiarum Hungaricae, 63, 327-341,, 2017, RefereedWebofScienceリポジトリ電子ジャーナル
  • An impressive new camaenid, Entadella entadiformis gen. & sp. n. from Guangxi, China (Gastropoda: Pulmonata)
    Páll-Gergely, B., Hunyadi, A., Otani, J.U. and Asami, T.
    Journal of Conchology, 42, 167-179,, 2016, RefereedWebofScience
  • Novel shell device for gas exchange in an operculate land snail
    Páll-Gergely, B., Naggs, F. and Asami, T.
    Biology Letters, 12, 20160151,, 2016, RefereedWebofScience電子ジャーナル
  • An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylommatophora: Helicarionidae)
    Páll-Gergely, B., Fehér, Z., Otani, J.U. and Asami, T.
    Molluscan Research, 37, 113-119,, 2016, RefereedWebofScience電子ジャーナル
  • A new species of Gudeodiscus Páll-Gergely, 2013 from China, with extraordinary conchological and anatomical features (Gastropoda, Pulmonata, Plectopylidae).
    Páll-Gergely, B. and Asami, T.
    Zookeys, 564, 1-19,, 2016, RefereedWebofScienceリポジトリ電子ジャーナル
  • A snail-eating snake recognizes prey-handedness
    Danaisawadi, P., Asami, T., Ota, H., Sutcharit, C. and Panha, S.
    SCIENTIFIC REPORTS, 6, 23832,, 2016, RefereedWebofScienceリポジトリ電子ジャーナル
  • The family Plectopylidae (Gastropoda, Pulmonata) in Laos with the description of two new genera and a new species
    Páll-Gergely, B., Muratov, I.V. and Asami, T.
    Zookeys, 592, 1-26,, 2016, RefereedWebofScienceリポジトリ電子ジャーナル
  • Predatory behavior of the snail-eating snake Pareas carinatus (Boie, 1828) (Squamata: Pareidae): an ethogram study
    Danaisawadi, P., Asami, T., Ota, H., Sutcharit, C. and Panha, S. [reprint]
    Tropical Natural History, 16, 21-31,, 2016, Refereedリポジトリ
  • A new Turkish species (Gastropoda: Pulmonata: Pristilomatidae) and association of distribution with habitat in the genus Vitrea
    Páll-Gergely, B. and Asami, T.
    Venus, 73, 41-50,, 2015, Refereed
  • Revision of the genera Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae)
    Páll-Gergely, B., Fehér, Z., Hunyadi, A. and Asami, T.
    Zootaxa, 3937, 1–49,, 2015, RefereedWebofScience電子ジャーナル
  • Subtle asymmetries in the snail-eating snake Pareas carinatus (Reptilia: Pareatidae)
    Danaisawadi, P., Asami, T., Ota, H., Sutcharit, C. and Panha, S.
    JOURNAL OF ETHOLOGY, 33, 243-246,, 2015, RefereedWebofScience電子ジャーナル
  • Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora).
    Páll-Gergely, B., Hunyadi, A., Ablett, J., Luong Van, H., Naggs, F. and Asami, T.
    Zookeys, 473, 1-118,, 2015, RefereedWebofScienceリポジトリ電子ジャーナル
  • A new Chinese species of Eostrobilops Pilsbry, 1927 with a checklist of Eostrobilops and Enteroplax Gude, 1897 species (Gastropoda, Pulmonata, Strobilopsidae)
    Páll-Gergely, B., Hunyadi, A. and Asami, T.
    Zookeys, 508, 85–95,, 2015, RefereedWebofScienceリポジトリ電子ジャーナル
  • Synchronous and non-synchronous semelparity in sibling species of pulmonates
    Nyumura, N. and Asami, T.
    Zoological Science, 32, 372-377,, 2015, RefereedWebofScienceリポジトリ電子ジャーナル
  • Seven new hypselostomatid species from China, including some of the world’s smallest land snails (Gastropoda, Pulmonata, Orthurethra)
    Páll-Gergely, B., Hunyadi, A., Jochum, A., and Asami, T.
    Zookeys, 523, 31–62,, 2015, RefereedWebofScienceリポジトリ電子ジャーナル
  • Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)
    Páll-Gergely, B., Budha, P.B., Naggs, F., Backeljau, T. and Asami, T.
    Zookeys, 529, 1–70,, 2015, RefereedWebofScienceリポジトリ電子ジャーナル
  • The contribution of seed dispersers to tree species diversity in tropical rain forests dispersers to tree species diversity in tropical rainforests
    Kakishima, S., Morita, S., Yoshida, K., Ishida, A., Hayashi, S., Asami, T., Ito, H., Miller III, D. G., Uehara, T., Mori, S., Hasegawa, E., Matsuura, K., Kasuya, E., Yoshimura, J.
    Royal Society Open Science, 2, 150330,, 2015, RefereedWebofScienceリポジトリ電子ジャーナル
  • Additional information on the distribution, anatomy, and systematics of living and fossil Chinese Plectopylidae (Gastropoda: Pulmonata)
    Páll-Gergely, B. and Asami, T. (543e39fe0cf25d6b1ad97280.pdf)
    Genus, 25, 527-564,, 2014, Refereed
  • Temperature is not responsible for left-right reversal in pelagic unicellular zooplanktons
    Ujiié, Y. and Asami, T.
    Journal of Zoology, 293, 16-24,, 2014, RefereedWebofScience電子ジャーナル
  • Evolutionary optimality in sex differences of longevity and athletic performances
    Asanuma, H., Kakisima, S., Ito, H., Kobayashi, K.,Hasegawa, E., Asami, T., Matsuura, K., Roff, D. A. and Yoshimura, J.
    SCIENTIFIC REPORTS, 4, 5425,, 2014, RefereedWebofScienceリポジトリ電子ジャーナル
  • Description of two new Ecuadorian Zilchistrophia Weyrauch 1960, with the clarification of the systematic position of the genus based on anatomical data (Gastropoda: Stylommatophora: Scolodontidae).
    Páll-Gergely, B. and Asami, T.
    Zookeys, 453, 1–17,, 2014, RefereedWebofScienceリポジトリ電子ジャーナル
  • A new, ribbed Schileykula species from north-eastern Turkey
    Páll-Gergely, B. and Asami, T.
    North-Western Journal of Zoology, 9, 214-216,, 2013, RefereedWebofScience電子ジャーナル
  • A peculiar new species in the genus Landouria Godwin-Austen, 1918 from China (Gastropoda: Heterobranchia: Stylommatophora: Camaenidae)
    Páll-Gergely, B, Hunyadi, A. and Asami, T.
    Molluscan Research, 33, 130–134,, 2013, RefereedWebofScience電子ジャーナル
  • Left-right asymmetric expression of dpp in the mantle of gastropods correlates with asymmetric shell coiling
    Shimizu, K., Iijima, M., Setiamarga, D. H. E., Sarashina, I., Kudoh, T., Asami, T., Gittenberger, E. and Endo, K.
    BMC EvoDevo, 4, 15, 2013, RefereedWebofScienceリポジトリ電子ジャーナル
  • Functional constraints on coiling geometry and aperture inclination in gastropods
    Noshita, K., Asami, T. and Ubukata, T.
    Paleobiology, 38, 322-334,, 2012, RefereedWebofScience電子ジャーナル
  • Longitudinal differentiation among pelagic populations in a planktic foraminifer
    Ujiié, Y., Asami, T., T. de Garidel-Thoron, Liu, H., Ishitani, Y. and De Vargas, C.
    Ecology and Evolution, 2, 1725–1737,, 2012, RefereedWebofScience電子ジャーナル
  • Chiral speciation in terrestrial pulmonate snails
    Gittenberger, E., Hamann, T. D. and Asami, T.
    PLOS ONE, 7, e34005,, 2012, RefereedWebofScienceリポジトリ電子ジャーナル
  • Internal selection against the evolutionof left-right reversal.
    Utsuno, H., Asami, T., Van Dooren, T. J. M. and Gittenberger, E.
    Evolution, 65, 2399–2411,, 2011, RefereedWebofScience電子ジャーナル
  • Genetic basis of racemism and ease of interchiral mating in a clausiliid species of snails.
    Utsuno, H., Kasem, S., Fukuda, H. and Asami, T.
    Molluscan Research, 30, 37-47,, 2010, RefereedWebofScience
  • Maternal inheritance of racemism in the terrestrial snail Bradybaena similaris.
    Utsuno, H. and Asami, T.
    JOURNAL OF HEREDITY, 101, 11-19,, 2010, RefereedWebofScience電子ジャーナル
  • Enantiomorphs differ in shape in opposite directions between populations.
    Nakadera, Y., Sutcharit, C., Ubukata, T., Seki, K., Utsuno, H., Panha, S. and Asami, T.
    Journal of Evolutionary Biology, 23, 2377-2384,, 2010, RefereedWebofScience電子ジャーナル
  • A speciation gene for left-right reversal in snails results in anti-predatoradaptation.
    Hoso, M., Kameda, Y., Wu, S.-P., Asami, T., Kato, M. and Hori, M.
    NATURE COMMUNICATIONS, 1, 133,, 2010, RefereedWebofScience電子ジャーナル
  • Population dynamics of Müllerian mimicry under interspecific competition.
    Kumazawa, F., Asami, T., Nakagiri, N., Tainaka, K., Togashi, T., Miyazaki, T., Yoshimura, J.
    Ecological Modelling, 220, 424-429,, 2009, RefereedWebofScience電子ジャーナル
  • Asymmetric reproductive isolation during simultaneous reciprocal mating in pulmonates.
    Wiwegweaw, A., Seki, K., Mori, H. and Asami, T.
    Biology Letters, 5, 240-243,, 2009, RefereedWebofScience電子ジャーナル
  • Fitness consequencesof reciprocally asymmetric hybridization between simultaneous hermaphrodites.
    Wiwegweaw, A., Seki, K.,Utsuno, H. and Asami, T.
    Zoological Science, 26, 191-196,, 2009, RefereedWebofScienceリポジトリ電子ジャーナル
  • Maintenance mechanism of a supergene for shell colour polymorphism in the terrestrial pulmonate Bradybaena similaris.
    Asami, T. and Asami, N.
    Basteria, 72, 119-127,, 2008, Refereed
  • Fibrous and helical calcite crystals induced by synthetic polypeptides containing 0-phospho-L-serine and 0-phospho-L-threonine
    Hayashi S, Ohkawa K, Suwa Y, Sugawara T, Asami T., and Yamamoto H.
    Macromolecular Bioscience, 8, 46-59,, 2008, RefereedWebofScience電子ジャーナル
  • Whole-body enantiomorphy and maternal inheritance of chiral reversal in the pond snail Lymnaea stagnalis.
    Asami, T., Gittenberger, E. and Falkner, G.
    JOURNAL OF HEREDITY, 99, 552-557,, 2008, RefereedWebofScience電子ジャーナル
  • Fluorescent pigment distinguishes between sibling snail species.
    Seki, K., Wiwegweaw, A. and Asami, T.
    Zoological Science, 25, 1212-1219,, 2008, RefereedWebofScience電子ジャーナル
  • The development and evolution of left-right asymmetry in invertebrates: lessons from Drosophila and snails.
    Okumura, T., Utsuno, H., Kuroda, J., Gittenberger, E., Asami, T. and Matsuno, K.
    Developmental Dynamics, 237, 3497-3515,, 2008, RefereedWebofScience電子ジャーナル
  • Polypolar spindle formation during the first cell cycle in the rainbow trout Oncorhynchus mykiss embryos after heat shock treatment
    Zhang, X., Asami, T. and Onozato H.
    Fishery Science, 73, 1325-1331,, 2007, RefereedWebofScience電子ジャーナル
  • Right-handed snakes: convergent evolution of asymmetry for functional specialization
    Hoso, M., Asami, T. and Hori, M.
    Biology Letters, 3, 169-172,, 2007, RefereedWebofScience電子ジャーナル
  • Evolution of whole-body enantiomorphy in the tree snail genus Amphidromus
    Sutcharit, C., Asami, T. and Panha, S.
    Journal of Evolutionary Biology, 20, 661-672,, 2007, RefereedWebofScience電子ジャーナル
  • Population dynamics of Batesian mimicry under interspecific competition
    Kumazawa, F., Asami, T., Hayashi, T. and Yoshimura, J.
    Evolutionary Ecology Research, 8, 591-604,, 2006, RefereedWebofScience
  • A geographical model of high species diversity.
    Tainaka, K., Itoh, Y., Yoshimura, J. and Asami, T.
    POPULATION ECOLOGY, 48, 113-119,, 2006, RefereedWebofScience電子ジャーナル
  • Lymnaea stagnalis as a model organism for studies of calcium carbonate biomineralization
    Endo, K., Sarashina, I., and Asami,T.
    Biomineralization: formation, diversity, evolution and application, 172-175, 2004, Refereed
  • Single-gene speciation by left-right reversal
    Ueshima, R. and Asami, T.
    Nature, 425, 679,, 2003, RefereedWebofScience電子ジャーナル
  • Geographical distributions of sibling species of land snails Bradybaena pellucida and B. similaris in the Boso Peninsula.
    Seki, K., Inoue, S. and Asami, T.
    Venus, 61, 41-48,, 2002, Refereed
  • Marine molluscan and brachiopod fauna of Tanoura, Nagashima Island, kaminoseki-cho, Yamaguchi Prefecture, Japan
    Fukuda, H., Asami, T., Yamashita, H., Sato, M, Hori, S. and Nakamura, Y.
    Yuriyagai, 7, 115-196,, 2001, Refereed
  • Effects of oviposition substrate on lifetime fecundity of the terrestrial pulmonate Bradybaena similaris
    Asami, T. and Ohbayashi, K.
    Journal of Conchology, 36, 1-9,, 1999, RefereedWebofScience
  • Evolution of mirror images by sexually asymmetric behavior in hermaphroditic snails
    Asami, T., Cowie, R. H. and Ohbayashi, K.
    American Naturalist, 152, 225-236,, 1998, RefereedWebofScience電子ジャーナル
  • Geographical distribution of the land snail Bradybaena pellucida (Pulmonata: Bradybaenidae)
    Asami, T., Yamashita, H., Park, J. and Ishikawa, H.
    Yuriyagai, 5, 31-42,, 1997, Refereed
  • The inheritance of shell color in the land snail Bradybaena pellucida
    Asami, T., Ohbayashi, K. and Seki, K.
    Venus, 56, 35-39,, 1997, Refereed電子ジャーナル
  • Melanism has not evolved in Japanese Biston betularia
    Asami, T. and Grant, B.
    Journal of the Lepidopterists'Society, 49, 88-91,, 1995, Refereed
  • A long term assessment of Biston betularia (L.) in one UK locality (Caldy Common, West Kirby, Wirral), 1959-93, and glimpses elsewhere.
    Clarke, C. A., Grant, B., Clarke, F. M. M. and Asami, T.
    Linnean, 10, 18-26,, 1994, Refereed
  • Genetic variation and evolution of coiling chirality in snails
    Asami, T.
    Forma, 9, 263-276,, 1993, Refereed
  • Divergence of activity patterns in sympatric land snails.
    Asami, T.
    Malacologia, 35, 399-406,, 1993, RefereedWebofScience
  • Interspecific differences in the tolerance of desiccation by juvenile land snails.
    Asami, T.
    Functional Ecology, 7, 571-577,, 1993, RefereedWebofScience電子ジャーナル
  • Inheritance of shell banding in the land snail Bradybaena pellusida.
    Asami, T., Fukuda, H. and Tomiyama, K.
    Venus, 52, 155-159,, 1993, Refereed電子ジャーナル
  • Temporal segregation of two sympatric species of land snails.
    Asami, T.
    Venus, 47, 278-297,, 1988, Refereed電子ジャーナル
  • The effect of maternal age upon fitness of progeny in the southern cowpea weevil, Callosobruchus maculatus.
    Wasserman, S. S. and Asami, T.
    Oikos, 45, 191-196,, 1985, Refereed電子ジャーナル
  • A simplified apparatus for vertical slab gel electrophoresis.
    Aotsuka, T. and Asami, T.
    Japanese Journal of Genetics, 54, 397-400,, 1979, Refereed電子ジャーナル

Lectures, oral presentations, etc.

  • What brings about left-right reversal in gastropods?
    Takahiro Asami
    Keynote, Southeast Asian Malacological Conference. online 15 July, 2022, Invited
  • Ecology and evolution of enantiomorphism in the tree snails Amphidromus
    Asami T, Sutcharit C, Panha S
    Frontiers in Taxonomy, International Conference on Biodiversity 2019, Bangkok, Thailand, 23 May, 2019, Invited
  • Handedness coevolution between snakes & snails in Southeast Asia
    Asami, T
    Keynote, The 4th Annual International Seminar on Trends in Science and Science Education 2017, Medan, Indonesia. 14 Nov, 2017, Faculty of Mathematics and Natural Sciences, State University of Medan, Invited
  • Evolution of reproductive barriers in snails
    Asami, T
    Keynote, World Congress of Malacology 2016, Penang, Malaysia. 23 Jul, 2016, Invited
  • Quantitative variation and evolution of chirality in pulmonate snails
    Asami, T
    Plenary Lecture, XI International Congress of Medical and Applied Malacology, Rio de Janeiro, Brazil. 25 Sep, 2012, Invited
  • Sex pheromones: Are hybrids superior or inferior in mating success?
    Asami, T
    Keynote, Symposium "Reproduction and Mating Systems in Hermaphroditic Molluscs" World Congress of Malacology 2010, Phuket, Thailand. 20 Jul, 2010, Invited

Research Themes

  • Evolution of chiral dimorphism in tree snails
    2024 - 2026
  • Quantitative asymmetry of spiral cleavage and reversal evolvability
    科学研究費補助金, Challenging Research (Exploratory), PI
    2019 - 2021
  • Chemical ecology of sexual isolation and reproductive interference in pulmonates
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
    2017 - 2021
  • Evolutionary biology of coexisting dynamics in snail chiral dimorphism
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
    2017 - 2021
  • Evolutionary ecology of developmental system: Why has situs inversus evolved only in internally fertilizing snails?
    科学研究費補助金, JSPS Grant-in-Aid for Challenging Exploratory Research, PI
    2014 - 2015
  • Developmental ecology for the animal rule of homochirality
    Grant-in-Aid for Scientific Research
  • Chemical ecology of hybrid vigor and hybrid breakdown in sex pheromone
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
    2012 - 2016
  • Ecological functions and long-term dynamics of snail chiral dimorphism
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
    2012 - 2016
  • Chemical ecology of sex phromones and premating isolation in hermaphrodites
    Frontier Research
  • Darwin 200: East Asian evolutionary diversity, JSPS Invitation Program for Asian Young Researchers, PI
    2009 - 2010
  • Dynamic equilibrium and adaptive evolution of snail chirality
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (B), PI
    2008 - 2011
  • Quantitative variation and adaptive evolution of left-right asymmetry
    Grant-in-Aid for Scientific Research
  • Handedness coevolution and ecological speciation
    Grant-in-Aid for Scientific Research
  • Asymmetric hybridization between species and molecular phylogeography of introgression
    Grant-in-Aid for Scientific Research
  • Maintenance mechanism and evolution of enantiomorphy in snails
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (B), PI
    2004 - 2007
  • Developmental constraint against left-right reversal
    科学研究費補助金, Grant-in-Aid for Exploratory Research, PI
    2004 - 2006
  • Evolutionary biology of enantiomorphy and mirror-image populations in tree snails
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (B), PI
    2001 - 2003
  • Is spiral cleavage responsible for morphogenetic handedness
    科学研究費補助金, JSPS Grant-in-Aid for Exploratory Research, PI
    2001 - 2002
  • Evolutionary ecology of mating behavior in enantiomorphic snails
    JST Basic Research Programs (Precursory Research for Embryonic Science and Technology :PRESTO)
  • Form and function of snail chirality, JST PRESTO, PI
    1999 - 2002
  • Pattern and process of chiral evolution in snails
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (C), PI
    1998 - 1999
  • Why snail are dextral: ecological genetics of chiral evolution, Kanagawa Academy of Science and Technology Research Grant, PI
    1998 - 1998
  • Chiral dimorphism and evolution in tree snails
    Grant-in-Aid for Scientific Research
  • Why snails are right-handed: ecological genetics of chiral reversal
    科学研究費補助金, JSPS Grant-in-Aid for Exploratory Research, PI
    1996 - 1997
  • Mechanisms of developmental constraint against chiral evolution
    Grant-in-Aid for Scientific Research
  • Interspecific interactions by allelopathy and predation in land snails
    科学研究費補助金, JSPS Grant-in-Aid for General Scientific Research (C), PI
    1994 - 1995
  • Why snails coil clockwise:Evolutionary exploration into coiling chirality, Narishige Zoology Grant, PI
    1994 - 1995
  • Comparative ecology of land-snail life history in suburbs, The Hino Green Fund, PI
    1993 - 1995
  • Interspecific interactions by allelopathy and predation, Sumitomo Grant for Basic Science Research Projects), PI
    1993 - 1994
  • Experimental methodology for promotion of scientific and cultural understandings of deleterious genetic variation and consanguineous-marriage tabooing, Nakayama Science Foundation Grant, PI.
    1993 - 1993
  • Educational use of genetic variation in wild populations: evolution and genetics by testing a hypothesis
    科学研究費補助金, JSPS Grant-in-Aid for Encouragement of Young Scientists (A), PI
    1992 - 1992
  • Allelopathy and multispecific interactions in terrestrial molluscs
    科学研究費補助金, JSPS Grant-in-Aid for Scientific Research on Priority Areas, PI
    1992 - 1992
  • Chiral variation and evolution
    Grant-in-Aid for Scientific Research
  • Educational use of genetic variation in wild populations: development of land-snail experiment
    科学研究費補助金, JSPS Grant-in-Aid for Encouragement of Young Scientists (A), PI
    1991 - 1991
  • Reproductive isolation mechanism
    Grant-in-Aid for Scientific Research
  • Speciation mechanism
    Grant-in-Aid for Scientific Research