Ph.D., University of Virginia

M.S., Tokyo Metropolitan University

Bachelor of Science, Tokyo Metropolitan University

Evolutionary biology, Ecological, population genetics

Ecology and environment, Evolutionary, behavioral, chemical ecology

Biodiversity and systematics, Phylogenetic systematics

2011
Professor, Department of Biology, Shinshu University

1999
Associate Professor, Department of Biology, Shinshu University

1995 - 1996
Field Museum of Natural History (Chicago), Bishop Museum (Honolulu)

1990
Smithonian Institute

1990
Associate Professor, Tokyo Metropolitan College

1989
Research Associate, School of Environmental Sciences, University of Virginia/ Virginia Museum of Natural History

1988, University of Virginia, School of Arts and Sciences, Biology

1982 - 1983, Rotary Scholar, University of Virginia

1978, Tokyo Metropolitan University, Faculty of Science, Biology

1980, Tokyo Metropolitan University, School of Sciences, Biology

2009
ZoologicalSociety of Japan, Zoological Science Award

Left–right reversal recurrently evolved regardless of diaphanous-related formin gene duplication or loss in snails
Takeshi Noda, Noriyuki Satoh, Edmund Gittenberger, Takahiro Asami
Journal of Molecular Evolution, 91, 721-729, 2023, Refereed

Two subspecies of the snail mite Riccardoella (Proriccardoella) reaumuri Fain & van Goethem, 1986 (Acari, Prostigmata, Ereynetidae) from Japan
Waki, T., Motochin, R., Asami, T., Shimano, S.
Systematic & Applied Acarology, 27, 839-854, 2022, Refereed

Genus-level revision of the Alycaeidae (Gastropoda, Cyclophoroidea), with an annotated species catalogue
Páll-Gergely, B., Sajan, S., Tripathy, B., Meng, K., Asami, T., Ablett, J. D.
Zookeys, 981, 1-220, 2020, Refereed

Riccardoella mites infecting six land snail species, referred to in the Red Data List and the Red Data Book
Waki T, Shimano S, Asami T
Taxa,, 45, 34-39, 2019, Refereed

Subspecies description rates are higher in morphologically complex land snails
Páll-Gergely B, Asami T, Sólymos P
Zoologica Scripta, 48, 185-193, 2019, Refereed

Evolution of a dextral lineage by left-right reversal in Cristataria (Gastropoda, Pulmonata, Clausiliidae)
Páll-Gergely P, Szekeres M, Fehér Z, Asami T, Harl J
Journal of Zoological Systematics and Evolutionary Research, 57, 520-526, 2019, Refereed

Three phyla—Two type specimens—One shell: History of a snail shell revealed by modern imaging technology
Schwaha T, Ruthensteiner B., Melzer R R, Asami T, Páll‐Gergely B
Journal of Zoological Systematics and Evolutionary Research, 57, 527-533, 2019, Refereed

Realtime Social Networking Service rapidly reveals distributions of non-indigenous land snails in a European capital
Páll-Gergely B, Majoros G, Domokos T, Juhász A, Turóci Á, Badacsonyi L, Fekete J, Asami T
BioInvasions Records, 8, 782–792, 2019, Refereed

Heterochirality results from reduction of maternal diaph expression in a terrestrial pulmonate snail
Noda T, Satoh N, Asami T
Zoology Letters, 5(https://doi.org/10.1186/s40851-018-0120-0), 2019, Refereed

Riccardoella reaumuri Fain and van Goethem, 1986 (Acariformes: Trombidiformes: Ereynetidae) from terrestrial mollusks in Okinawajima Island, as first record from Japan
Waki T, Shimano S, Asami T, Miyai T, Sasaki T
The biological magazine Okinawa, 56, 27-31, 2018, Refereed

Enantiomorphs and taxonomy of three conchological species in flat-shelled snails Trichocathaica (Pulmonata: Camaenidae)
Páll-Gergely B, Hunyadi A, Asami T
Zookeys, 810, 19–44, 2018, Refereed

Taxonomical notes on Chinese camaenids with description of three new species (Gastropoda: Pulmonata)
Wu, M, Asami, T
Molluscan Research, 38, 137-148,, 2018, Refereed

A new species of Camaenidae (Gastropoda, Pulmonata) from Nusa Penida and Lombok Islands, Indonesia: novelty in a well-known fauna
Páll-Gergely, B, Otani, JU, Hosoda, T, Asami, T, Harl, J
Molluscan Research, 38, 41-49,, 2018, Refereed

A new species of Metalycaeus hitherto undistinguished from M. vinctus (Pilsbry, 1902) (Gastropoda: Cyclophoroidea: Alycaeidae)
Páll-Gergely, B, Asami, T
Venus, 75, 1–16,, 2017, Refereed

Molecular systematics of the land snail family Orculidae reveal paraphyly and deep splits within the clade Orthurethra (Gastropoda: Pulmonata)
Harl, J, Haring, E, Sittenthaler, M, Sattmann, H, Asami, T, Páll-Gergely, B
Zoological Journal of the Linnean Society, 181, 778–794,, 2017, Refereed

Revision of the Alycaeidae of China, Laos and Vietnam (Gastropoda: Cyclophoroidea) I: The genera Dicharax and Metalycaeus
Páll-Gergely, B, Hunyadi, A, Đỗ, Đ S, Naggs, F, Asami, T
Zootaxa,, 4331, 1–124, 2017, Refereed

Breathing device of a new Streptaulus species from Vietnam extends understanding of the function and structure of respiratory tubes in cyclophoroids (Gastropoda: Caenogastropoda: Pupinidae)
Páll-Gergely, B, Gargominy, O, Fontaine, B, Asami, T
Journal of Molluscan Studies, 83, 243-248,, 2017, Refereed

A new diplommatinid genus and two new species from the Philippines (Gastropoda, Caenogastropoda, Cyclophoroidea)
Páll-Gergely, B, Hunyadi, A, Asami, T
Zookeys, 678, 1-10,, 2017, Refereed

Three new species and a new genus of Hypselostomatidae (Gastropoda: Pulmonata) from Cong Troi Cave, Northern Vietnam
Páll-Gergely B, Jochum, A, Asami, T
Acta Zoologica Academiae Scientiarum Hungaricae, 63, 327-341,, 2017, Refereed

An impressive new camaenid, Entadella entadiformis gen. & sp. n. from Guangxi, China (Gastropoda: Pulmonata)
Páll-Gergely, B., Hunyadi, A., Otani, J.U. and Asami, T.
Journal of Conchology, 42, 167-179,, 2016, Refereed

Novel shell device for gas exchange in an operculate land snail
Páll-Gergely, B., Naggs, F. and Asami, T.
Biology Letters, 12, 20160151,, 2016, Refereed

An integrative taxonomic approach to infer the systematic position of Chalepotaxis Ancey, 1887 (Gastropoda: Stylommatophora: Helicarionidae)
Páll-Gergely, B., Fehér, Z., Otani, J.U. and Asami, T.
Molluscan Research, 37, 113-119,, 2016, Refereed

A new species of Gudeodiscus Páll-Gergely, 2013 from China, with extraordinary conchological and anatomical features (Gastropoda, Pulmonata, Plectopylidae).
Páll-Gergely, B. and Asami, T.
Zookeys, 564, 1-19,, 2016, Refereed

A snail-eating snake recognizes prey-handedness
Danaisawadi, P., Asami, T., Ota, H., Sutcharit, C. and Panha, S.
SCIENTIFIC REPORTS, 6, 23832,, 2016, Refereed

The family Plectopylidae (Gastropoda, Pulmonata) in Laos with the description of two new genera and a new species
Páll-Gergely, B., Muratov, I.V. and Asami, T.
Zookeys, 592, 1-26,, 2016, Refereed

Predatory behavior of the snail-eating snake Pareas carinatus (Boie, 1828) (Squamata: Pareidae): an ethogram study
Danaisawadi, P., Asami, T., Ota, H., Sutcharit, C. and Panha, S. [reprint]
Tropical Natural History, 16, 21-31,, 2016, Refereed

A new Turkish species (Gastropoda: Pulmonata: Pristilomatidae) and association of distribution with habitat in the genus Vitrea
Páll-Gergely, B. and Asami, T. https://www.jstage.jst.go.jp/article/venus/73/1-2/73_41/_pdf
Venus, 73, 41-50,, 2015, Refereed

Revision of the genera Pseudopomatias and its relatives (Gastropoda: Cyclophoroidea: Pupinidae)
Páll-Gergely, B., Fehér, Z., Hunyadi, A. and Asami, T.
Zootaxa, 3937, 1–49,, 2015, Refereed

Subtle asymmetries in the snail-eating snake Pareas carinatus (Reptilia: Pareatidae)
Danaisawadi, P., Asami, T., Ota, H., Sutcharit, C. and Panha, S.
JOURNAL OF ETHOLOGY, 33, 243-246,, 2015, Refereed

Systematics of the family Plectopylidae in Vietnam with additional information on Chinese taxa (Gastropoda, Pulmonata, Stylommatophora).
Páll-Gergely, B., Hunyadi, A., Ablett, J., Luong Van, H., Naggs, F. and Asami, T.
Zookeys, 473, 1-118,, 2015, Refereed

A new Chinese species of Eostrobilops Pilsbry, 1927 with a checklist of Eostrobilops and Enteroplax Gude, 1897 species (Gastropoda, Pulmonata, Strobilopsidae)
Páll-Gergely, B., Hunyadi, A. and Asami, T.
Zookeys, 508, 85–95,, 2015, Refereed

Synchronous and non-synchronous semelparity in sibling species of pulmonates
Nyumura, N. and Asami, T.
Zoological Science, 32, 372-377,, 2015, Refereed

Seven new hypselostomatid species from China, including some of the world’s smallest land snails (Gastropoda, Pulmonata, Orthurethra)
Páll-Gergely, B., Hunyadi, A., Jochum, A., and Asami, T.
Zookeys, 523, 31–62,, 2015, Refereed

Review of the genus Endothyrella Zilch, 1960 with description of five new species (Gastropoda, Pulmonata, Plectopylidae)
Páll-Gergely, B., Budha, P.B., Naggs, F., Backeljau, T. and Asami, T.
Zookeys, 529, 1–70,, 2015, Refereed

The contribution of seed dispersers to tree species diversity in tropical rain forests dispersers to tree species diversity in tropical rainforests
Kakishima, S., Morita, S., Yoshida, K., Ishida, A., Hayashi, S., Asami, T., Ito, H., Miller III, D. G., Uehara, T., Mori, S., Hasegawa, E., Matsuura, K., Kasuya, E., Yoshimura, J.
Royal Society Open Science, 2, 150330,, 2015, Refereed

Additional information on the distribution, anatomy, and systematics of living and fossil Chinese Plectopylidae (Gastropoda: Pulmonata)
Páll-Gergely, B. and Asami, T. (543e39fe0cf25d6b1ad97280.pdf)
Genus, 25, 527-564,, 2014, Refereed

Temperature is not responsible for left-right reversal in pelagic unicellular zooplanktons
Ujiié, Y. and Asami, T.
Journal of Zoology, 293, 16-24,, 2014, Refereed

Evolutionary optimality in sex differences of longevity and athletic performances
Asanuma, H., Kakisima, S., Ito, H., Kobayashi, K.,Hasegawa, E., Asami, T., Matsuura, K., Roff, D. A. and Yoshimura, J.
SCIENTIFIC REPORTS, 4, 5425,, 2014, Refereed

Description of two new Ecuadorian Zilchistrophia Weyrauch 1960, with the clarification of the systematic position of the genus based on anatomical data (Gastropoda: Stylommatophora: Scolodontidae).
Páll-Gergely, B. and Asami, T.
Zookeys, 453, 1–17,, 2014, Refereed

A new, ribbed Schileykula species from north-eastern Turkey
Páll-Gergely, B. and Asami, T. http://biozoojournals.ro/nwjz/content/v9n1/nwjz.132304.Pall-Gergely.pdf
North-Western Journal of Zoology, 9, 214-216,, 2013, Refereed

A peculiar new species in the genus Landouria Godwin-Austen, 1918 from China (Gastropoda: Heterobranchia: Stylommatophora: Camaenidae)
Páll-Gergely, B, Hunyadi, A. and Asami, T.
Molluscan Research, 33, 130–134,, 2013, Refereed

Left-right asymmetric expression of dpp in the mantle of gastropods correlates with asymmetric shell coiling
Shimizu, K., Iijima, M., Setiamarga, D. H. E., Sarashina, I., Kudoh, T., Asami, T., Gittenberger, E. and Endo, K.
BMC EvoDevo, 4, 15, 2013, Refereed

Functional constraints on coiling geometry and aperture inclination in gastropods
Noshita, K., Asami, T. and Ubukata, T.
Paleobiology, 38, 322-334,, 2012, Refereed

Longitudinal differentiation among pelagic populations in a planktic foraminifer
Ujiié, Y., Asami, T., T. de Garidel-Thoron, Liu, H., Ishitani, Y. and De Vargas, C.
Ecology and Evolution, 2, 1725–1737,, 2012, Refereed

Chiral speciation in terrestrial pulmonate snails
Gittenberger, E., Hamann, T. D. and Asami, T.
PLOS ONE, 7, e34005,, 2012, Refereed

Internal selection against the evolutionof left-right reversal.
Utsuno, H., Asami, T., Van Dooren, T. J. M. and Gittenberger, E.
Evolution, 65, 2399–2411,, 2011, Refereed

Genetic basis of racemism and ease of interchiral mating in a clausiliid species of snails.
Utsuno, H., Kasem, S., Fukuda, H. and Asami, T.
Molluscan Research, 30, 37-47,, 2010, Refereed

Maternal inheritance of racemism in the terrestrial snail Bradybaena similaris.
Utsuno, H. and Asami, T.
JOURNAL OF HEREDITY, 101, 11-19,, 2010, Refereed

Enantiomorphs differ in shape in opposite directions between populations.
Nakadera, Y., Sutcharit, C., Ubukata, T., Seki, K., Utsuno, H., Panha, S. and Asami, T.
Journal of Evolutionary Biology, 23, 2377-2384,, 2010, Refereed

A speciation gene for left-right reversal in snails results in anti-predatoradaptation.
Hoso, M., Kameda, Y., Wu, S.-P., Asami, T., Kato, M. and Hori, M.
NATURE COMMUNICATIONS, 1, 133,, 2010, Refereed

Population dynamics of Müllerian mimicry under interspecific competition.
Kumazawa, F., Asami, T., Nakagiri, N., Tainaka, K., Togashi, T., Miyazaki, T., Yoshimura, J.
Ecological Modelling, 220, 424-429,, 2009, Refereed

Asymmetric reproductive isolation during simultaneous reciprocal mating in pulmonates.
Wiwegweaw, A., Seki, K., Mori, H. and Asami, T.
Biology Letters, 5, 240-243,, 2009, Refereed

Fitness consequencesof reciprocally asymmetric hybridization between simultaneous hermaphrodites.
Wiwegweaw, A., Seki, K.,Utsuno, H. and Asami, T.
Zoological Science, 26, 191-196,, 2009, Refereed

Maintenance mechanism of a supergene for shell colour polymorphism in the terrestrial pulmonate Bradybaena similaris.
Asami, T. and Asami, N.
Basteria, 72, 119-127,, 2008, Refereed

Fibrous and helical calcite crystals induced by synthetic polypeptides containing 0-phospho-L-serine and 0-phospho-L-threonine
Hayashi S, Ohkawa K, Suwa Y, Sugawara T, Asami T., and Yamamoto H.
Macromolecular Bioscience, 8, 46-59,, 2008, Refereed

Whole-body enantiomorphy and maternal inheritance of chiral reversal in the pond snail Lymnaea stagnalis.
Asami, T., Gittenberger, E. and Falkner, G.
JOURNAL OF HEREDITY, 99, 552-557,, 2008, Refereed

Fluorescent pigment distinguishes between sibling snail species.
Seki, K., Wiwegweaw, A. and Asami, T.
Zoological Science, 25, 1212-1219,, 2008, Refereed

The development and evolution of left-right asymmetry in invertebrates: lessons from Drosophila and snails.
Okumura, T., Utsuno, H., Kuroda, J., Gittenberger, E., Asami, T. and Matsuno, K.
Developmental Dynamics, 237, 3497-3515,, 2008, Refereed

Polypolar spindle formation during the first cell cycle in the rainbow trout Oncorhynchus mykiss embryos after heat shock treatment
Zhang, X., Asami, T. and Onozato H.
Fishery Science, 73, 1325-1331,, 2007, Refereed

Right-handed snakes: convergent evolution of asymmetry for functional specialization
Hoso, M., Asami, T. and Hori, M.
Biology Letters, 3, 169-172,, 2007, Refereed

Evolution of whole-body enantiomorphy in the tree snail genus Amphidromus
Sutcharit, C., Asami, T. and Panha, S.
Journal of Evolutionary Biology, 20, 661-672,, 2007, Refereed

Population dynamics of Batesian mimicry under interspecific competition
Kumazawa, F., Asami, T., Hayashi, T. and Yoshimura, J.
Evolutionary Ecology Research, 8, 591-604,, 2006, Refereed

A geographical model of high species diversity.
Tainaka, K., Itoh, Y., Yoshimura, J. and Asami, T.
POPULATION ECOLOGY, 48, 113-119,, 2006, Refereed

Lymnaea stagnalis as a model organism for studies of calcium carbonate biomineralization
Endo, K., Sarashina, I., and Asami,T.
Biomineralization: formation, diversity, evolution and application, 172-175, 2004, Refereed

Single-gene speciation by left-right reversal
Ueshima, R. and Asami, T.
Nature, 425, 679,, 2003, Refereed

Geographical distributions of sibling species of land snails Bradybaena pellucida and B. similaris in the Boso Peninsula.
Seki, K., Inoue, S. and Asami, T.
Venus, 61, 41-48,, 2002, Refereed

Marine molluscan and brachiopod fauna of Tanoura, Nagashima Island, kaminoseki-cho, Yamaguchi Prefecture, Japan
Fukuda, H., Asami, T., Yamashita, H., Sato, M, Hori, S. and Nakamura, Y.
Yuriyagai, 7, 115-196,, 2001, Refereed

Effects of oviposition substrate on lifetime fecundity of the terrestrial pulmonate Bradybaena similaris
Asami, T. and Ohbayashi, K.
Journal of Conchology, 36, 1-9,, 1999, Refereed

Evolution of mirror images by sexually asymmetric behavior in hermaphroditic snails
Asami, T., Cowie, R. H. and Ohbayashi, K.
American Naturalist, 152, 225-236,, 1998, Refereed

Geographical distribution of the land snail Bradybaena pellucida (Pulmonata: Bradybaenidae)
Asami, T., Yamashita, H., Park, J. and Ishikawa, H.
Yuriyagai, 5, 31-42,, 1997, Refereed

The inheritance of shell color in the land snail Bradybaena pellucida
Asami, T., Ohbayashi, K. and Seki, K.
Venus, 56, 35-39,, 1997, Refereed

Melanism has not evolved in Japanese Biston betularia
Asami, T. and Grant, B.
Journal of the Lepidopterists'Society, 49, 88-91,, 1995, Refereed

A long term assessment of Biston betularia (L.) in one UK locality (Caldy Common, West Kirby, Wirral), 1959-93, and glimpses elsewhere.
Clarke, C. A., Grant, B., Clarke, F. M. M. and Asami, T.
Linnean, 10, 18-26,, 1994, Refereed

Genetic variation and evolution of coiling chirality in snails
Asami, T. http://www.scipress.org/journals/forma/abstract/0802/08020263.html
Forma, 9, 263-276,, 1993, Refereed

Divergence of activity patterns in sympatric land snails.
Asami, T.
Malacologia, 35, 399-406,, 1993, Refereed

Interspecific differences in the tolerance of desiccation by juvenile land snails.
Asami, T.
Functional Ecology, 7, 571-577,, 1993, Refereed

Inheritance of shell banding in the land snail Bradybaena pellusida.
Asami, T., Fukuda, H. and Tomiyama, K.
Venus, 52, 155-159,, 1993, Refereed

Temporal segregation of two sympatric species of land snails.
Asami, T.
Venus, 47, 278-297,, 1988, Refereed

The effect of maternal age upon fitness of progeny in the southern cowpea weevil, Callosobruchus maculatus.
Wasserman, S. S. and Asami, T.
Oikos, 45, 191-196,, 1985, Refereed

A simplified apparatus for vertical slab gel electrophoresis.
Aotsuka, T. and Asami, T. https://www.jstage.jst.go.jp/article/ggs1921/54/5/54_5_397/_pdf
Japanese Journal of Genetics, 54, 397-400,, 1979, Refereed

What brings about left-right reversal in gastropods?
Takahiro Asami
Keynote, Southeast Asian Malacological Conference. online 15 July, 2022, Invited

Ecology and evolution of enantiomorphism in the tree snails Amphidromus
Asami T, Sutcharit C, Panha S
Frontiers in Taxonomy, International Conference on Biodiversity 2019, Bangkok, Thailand, 23 May, 2019, Invited

Handedness coevolution between snakes & snails in Southeast Asia
Asami, T
Keynote, The 4th Annual International Seminar on Trends in Science and Science Education 2017, Medan, Indonesia. 14 Nov, 2017, Faculty of Mathematics and Natural Sciences, State University of Medan, Invited

Evolution of reproductive barriers in snails
Asami, T
Keynote, World Congress of Malacology 2016, Penang, Malaysia. 23 Jul, 2016, Invited

Quantitative variation and evolution of chirality in pulmonate snails
Asami, T
Plenary Lecture, XI International Congress of Medical and Applied Malacology, Rio de Janeiro, Brazil. 25 Sep, 2012, Invited

Sex pheromones: Are hybrids superior or inferior in mating success?
Asami, T
Keynote, Symposium "Reproduction and Mating Systems in Hermaphroditic Molluscs" World Congress of Malacology 2010, Phuket, Thailand. 20 Jul, 2010, Invited

Evolution of chiral dimorphism in tree snails
Grants-in-Aid for Scientific Research, Japan Society for the Promotion of Science (JSPS), Grant-in-Aid for Scientific Research (C)
2024 - 2026

Quantitative asymmetry of spiral cleavage and reversal evolvability
科学研究費補助金, Challenging Research (Exploratory), PI
2019 - 2021

Chemical ecology of sexual isolation and reproductive interference in pulmonates
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
2017 - 2021

Evolutionary biology of coexisting dynamics in snail chiral dimorphism
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
2017 - 2021

Evolutionary ecology of developmental system: Why has situs inversus evolved only in internally fertilizing snails?
科学研究費補助金, JSPS Grant-in-Aid for Challenging Exploratory Research, PI
2014 - 2015

Chemical ecology of hybrid vigor and hybrid breakdown in sex pheromone
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
2012 - 2016

Ecological functions and long-term dynamics of snail chiral dimorphism
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (A), PI
2012 - 2016

Darwin 200: East Asian evolutionary diversity, JSPS Invitation Program for Asian Young Researchers, PI
2009 - 2010

Dynamic equilibrium and adaptive evolution of snail chirality
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (B), PI
2008 - 2011

Maintenance mechanism and evolution of enantiomorphy in snails
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (B), PI
2004 - 2007

Developmental constraint against left-right reversal
科学研究費補助金, Grant-in-Aid for Exploratory Research, PI
2004 - 2006

Evolutionary biology of enantiomorphy and mirror-image populations in tree snails
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (B), PI
2001 - 2003

Is spiral cleavage responsible for morphogenetic handedness
科学研究費補助金, JSPS Grant-in-Aid for Exploratory Research, PI
2001 - 2002

Form and function of snail chirality, JST PRESTO, PI
1999 - 2002

Pattern and process of chiral evolution in snails
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research (C), PI
1998 - 1999

Why snail are dextral: ecological genetics of chiral evolution, Kanagawa Academy of Science and Technology Research Grant, PI
1998 - 1998

Why snails are right-handed: ecological genetics of chiral reversal
科学研究費補助金, JSPS Grant-in-Aid for Exploratory Research, PI
1996 - 1997

Interspecific interactions by allelopathy and predation in land snails
科学研究費補助金, JSPS Grant-in-Aid for General Scientific Research (C), PI
1994 - 1995

Why snails coil clockwise:Evolutionary exploration into coiling chirality, Narishige Zoology Grant, PI
1994 - 1995

Comparative ecology of land-snail life history in suburbs, The Hino Green Fund, PI
1993 - 1995

Interspecific interactions by allelopathy and predation, Sumitomo Grant for Basic Science Research Projects), PI
1993 - 1994

Experimental methodology for promotion of scientific and cultural understandings of deleterious genetic variation and consanguineous-marriage tabooing, Nakayama Science Foundation Grant, PI.
1993 - 1993

Educational use of genetic variation in wild populations: evolution and genetics by testing a hypothesis
科学研究費補助金, JSPS Grant-in-Aid for Encouragement of Young Scientists (A), PI
1992 - 1992

Allelopathy and multispecific interactions in terrestrial molluscs
科学研究費補助金, JSPS Grant-in-Aid for Scientific Research on Priority Areas, PI
1992 - 1992

Educational use of genetic variation in wild populations: development of land-snail experiment
科学研究費補助金, JSPS Grant-in-Aid for Encouragement of Young Scientists (A), PI
1991 - 1991