Developmental biology

Ecology and environment

Biodiversity and systematics

Evolutionary biology

2022
Faculty of Science Department of Science, Shinshu University

2021 - 2022
Faculty of Life and Environmental Sciences, University of Tsukuba

2019 - 2021
National Institute for Basic Biology

2016 - 2019
Interdisciplinary Graduate School of Science and Technology, Shinshu University

2016 - 2019, Shinshu University, Interdisciplinary Graduate School of Science and Technology

2023
河川財団, 河川財団奨励賞
水生昆虫における正確なDNAバーコディング領域の解析手法の確立と実証, International academic award
竹中將起

2022
日本陸水学会, 国際交流奨励賞
International academic award
竹中將起

2020
日本動物学会, 日本動物学会 藤井賞
International academic award
竹中將起・関根一希・東城幸治

2020
Zoological Society of Japan, 2019, Zoological Science Award
The First Establishment of “Hand-Pairing” Cross-Breeding Method for the Most Ancestral Wing Acquired Insect Group Masaki Takenaka, Kazuki Sekiné, Koji Tojo, International academic award
Masaki Takenaka;Kazuki Sekiné;Koji Tojo

2019
河川基金, 河川基金優秀成果
千曲-信濃川水系で新規発見されたカワヨシノボリ集団の遺伝構造：その起源の究明と保全, International academic award
竹中將起

Environmental DNA metabarcoding on aquatic insects: Comparing the primer sets of MtInsects‐16S based on the mtDNA 16S and general marker based on the mtDNA COI region
Masaki Takenaka; Yuta Hasebe; Koki Yano; Seiya Okamoto; Koji Tojo; Masashi Seki; Shun Sekiguchi; Takeshi Jitsumasa; Nobuyuki Morohashi; Yoshihiro Handa; Tomoka Sakaba
Environmental DNA, 6(4), Jul. 2024, Refereed
Lead, Corresponding

Three new species of the genera Ceraclea Stephens and Setodes Rambur (Trichoptera: Leptoceridae) from Laos and Cambodia
PONGSAK LAUDEE; HANS MALICKY; MASAKI TAKENAKA; NANNAPHAT SUWANNARAT; CHAMROEUN KONG; CHANDA VONGSWOMBATH; KOJI TOJO
Zootaxa, 5471(1), 134-140, Jun. 2024, Refereed
Lead

Evolutionary history of a cold-adapted limnephilid caddisfly: Effects of climate change and topography on genetic structure
Hirohisa Suzuki; Masaki Takenaka; Koji Tojo
Molecular Phylogenetics and Evolution, 191, 107967-107967, Feb. 2024, Refereed, Not invited
Lead

Variations in the phenological patterns of a caddisfly inhabiting the same mountain massifs: life history differences in different altitudinal zones
Suzuki H; Takenaka M; Tojo K
Ecology and Evolution, accepted, 2024, Refereed
Lead

Redescription of Utaperla orientalis (Plecoptera, Chroloperlidae) from Hokkaido: The first record of a Utaperla stonefly in Japan
Yoshinari G; Inada K; Takenaka M; Tojo K; Negishi J
Limnology, accepted, 2024, Refereed
Lead

Phylogeography of an insect inhabiting ‘Sky Islands’: the relationships among genetic structures and geographical characteristics, geohistorical characteristics, and cyclical climate changes
Hirohisa Suzuki; Masaki Takenaka; Koji Tojo
Biological Journal of the Linnean Society, Nov. 2023, Refereed, Not invited
Lead

New species of the genus Trichosetodes Ulmer, 1915 (Trichoptera: Leptoceridae) from Ratanakiri province, Cambodia, based on morphological and molecular data
LAUDEE P; MALICKY P; KONG C; TAKENAKA M; TOJO K
Zookeys, 1182, 153-164, Oct. 2023, Refereed
Lead

Phylogeography of the true freshwater crab, Geothelphusa dehaani: detected dual dispersal routes via land and sea
Takenaka Masaki; Yano Koki; Tojo Koji
Zoology, 160, 126118-126118, Aug. 2023, Refereed
Lead

Molecular Phylogeography of Japanese Long-Toed Water Beetles, Elmomorphus (Coleoptera: Dryopoidea: Dryopidae): First Record of Elmomorphus amamiensis from Okinoerabu-jima Island
Takumi YOSHIDA; Masaki TAKENAKA; Koji TOJO
Japanese Journal of Entomology (New Series), 26, 165-174, Aug. 2023, Refereed
Lead

Phylogeography of alpine and subalpine adapted Pseudostenophylax caddisflies (Limnephilidae: Trichoptera): a strong relationship with mountain formation
Mikami K; Takenaka M; Nozaki T; Bae YJ; Tojo K
Biological Journal of the Linnean Society, Jun. 2023, Refereed
Lead

Ephemeropteran fauna of the Koshiki Islands
吉田匠; 野田聖; 東城幸治; 竹中將起
昆蟲（ニューシリーズ）, 26, 107-114, Jun. 2023, Refereed
Corresponding

Genetic differentiation within species exhibiting widespread gene flow; phylogeography of the downstream-inhabiting speciesEphemera orientalis(Insecta: Ephemeroptera)
Masaki Takenaka; Hiroaki Kogawara; Yeon Jae Bae; Koji Tojo
Biological Journal of the Linnean Society, 138(4), 351-364, Mar. 2023, Refereed
Lead

Development of novel PCR primer sets for DNA barcoding of aquatic insects, and the discovery of some cryptic species
Masaki Takenaka; Koki Yano; Tomoya Suzuki; Koji Tojo
Limnology, 24(2), 121-136, Jan. 2023, Refereed
Lead, Corresponding

Behavior of snow monkeys hunting fish to survive winter
Takenaka, M; Hayashi, K; Yamada, G; Ogura, T; Ito, M; Milner, A.M; Tojo, K
Scientific Reports, 12(20324), Dec. 2022, Refereed, Not invited
Lead, Corresponding

Seasonal modifications of longitudinal distribution patterns within a stream: Interspecific interactions in the niche overlap zones of two Ephemera mayflies
Seiya Okamoto; Masaki Takenaka; Koji Tojo
Ecology and Evolution, 12(e8766), 2022, Refereed
Lead

Winter diet of Japanese macaques from Chubu Sangaku National Park, Japan incorporates freshwater biota
Alexander M. Milner; Susanna A. Wood; Catherine Docherty; Laura Biessy; Masaki Takenaka; Koji Tojo
Scientific Reports, 11(1), Dec. 2021, Refereed
Lead

Phylogeography of the northernmost distributed Anisocentropus caddisflies and their comparative genetic structures based on habitat preferences
Masaki Takenaka; Saki Shibata; Tomiko Ito; Noriyoshi Shimura; Koji Tojo
Ecology and Evolution, 11(9), 4957-4971, May 2021, Refereed, Not invited
Lead

Wide-Scale Gene Flow, Even in Insects that have Lost their Flight Ability: Presence of Dispersion Due to a Unique Parasitic Ecological Strategy of Piggybacking Hosts
Ohjiro Ohnishi; Masaki Takenaka; Ryosuke Okano; Hiroyuki Yoshitomi; Koji Tojo
Zoological Science, 38(2), Jan. 2021, Refereed
Lead

北海道札内川・戸蔦別川で採集されたカワゲラ類
https; doi.org; nlandwaterbiology
陸水生物学報, 36, 19-28, 2021, Refereed, Not invited
Lead

Identifying a “pseudogene” for the mitochondrial DNA COI region of the corixid aquatic insect, Hesperocorixa distanti (Heteroptera, Corixidae)
Koki Yano; Masaki Takenaka; Toshimasa Mitamura; Koji Tojo
Limnology, 21(3), 319-325, Aug. 2020, Refereed
Lead

Development of microsatellite markers for a giant water bug, Appasus japonicus, distributed in East Asia
Tomoya Suzuki; Akira S. Hirao; Masaki Takenaka; Koki Yano; Koji Tojo
Genes & Genetic Systems, 95, 323-329, May 2020, Refereed
Lead

Genealogical Position of Japanese Populations of the Globally Distributed Mayfly Cloeon dipterum and Related Species (Ephemeroptera, Baetidae): A Molecular Phylogeographic Analysis
Koki Yano; Masaki Takenaka; Koji Tojo
Zoological Science, 36(6), 479-479, Dec. 2019, Refereed
Lead

Concordance between molecular biogeography of Dipteromimus tipuliformis and geological history in the local fine scale (Ephemeroptera, Dipteromimidae)
Masaki Takenaka; Tetsuya Tokiwa; Koji Tojo
Molecular Phylogenetics and Evolution, 139, 106547-106547, Oct. 2019, Refereed
Lead

The First Establishment of “Hand-Pairing” Cross-Breeding Method for the Most Ancestral Wing Acquired Insect Group
Masaki Takenaka; Kazuki Sekiné; Koji Tojo
Zoological Science, 36(2), 136-136, Apr. 2019, Refereed
Lead

Ancient origin of a dipteromimid mayfly family endemic to the Japanese Islands and its genetic differentiation across tectonic faults
Takenaka Masaki; Tojo Koji
BIOLOGICAL JOURNAL OF THE LINNEAN SOCIETY, 126(3), 555-573, Mar. 2019, Refereed, Not invited
Lead

Evaluation of the Effectiveness of a “Hand-Pairing” Cross-Breeding Experiment in Ephemeroptera
Takenaka Masaki; Sekine Kazuki; Yano Koki; Tojo Koji
Zoosymposia, 16, 210-217, 2019, Refereed, Not invited
Lead

Update on distribution area information of three species of Anisocentropus caddisflies Inhabiting the Japanese Islands (Trichoptera: Calamoceratidae)
Takenaka, M; S. Shibata; N. Shimura; T. Ito; K. Tojo
New Entomologist, 67, 13-20, 2018, Refereed, Not invited
Lead

Species diversity of insects in Japan: Their origins and diversification processes
Koji Tojo; Kazuki Sekiné; Masaki Takenaka; Yuichi Isaka; Shohei Komaki; Tomoya Suzuki; Sean D. Schoville
Entomological Science, 20(1), 357-381, Jan. 2017, Refereed, Invited
Lead

First Record of the Ephemerid May y Ephemera formosana from the Tokashiki-jima Island, of the Ryukyu Archipelago
竹中將起; 趙在翼; 東城幸治
New Entomologist, 66, 30-32, 2017, Refereed, Not invited
Lead

Record of a Giant Water Bug, Lethocerus deyrollei, in Joso City (Heteroptera, Belostomatidae)
東城幸治; 竹中將起
New Entomologist, 65, 122-123, 2016, Refereed, Not invited
Lead

Light Trap Collection of Two Mayfly Species, Ephoron eophilum and Bracycercus japonicus, after the Kinu-gawa River Flood of September 2015
東城幸治; 竹中將起
New Entomologist, 65, 119-121, 2016, Refereed, Not invited
Lead

A study into the distribution of the family Dipteromimidae, which is endemic to the Japanese Islands
Takenaka, M; K. Tojo
Biol. Inl. Wat., Suppl. 3, 93-97., 2016, Refereed, Not invited
Lead

The Record of at the Site of the Most Head water Region of Pseudostenophylax ondakensis the Yari-sawa Stream on Mt.Yari-gatake
竹中將起; 東城幸治
New Entomologist, 64, 60-62, 2015, Refereed, Not invited
Lead

The First Record of thePeltoperlid Stonefly, Cryptoperla japonica on Sado Island
竹中將起; 斎藤梨絵; 鈴木智也; 大西央士郎; 富樫繁春; 岸本亨; 東城幸治
New entomologist, 64(1-2), 19-23, 2015, Refereed, Not invited
Lead

The Distribution Range of the Dipteromimid Mayfles (Ephemeroptera, Dipteromimidae)
竹中將起; 小河原寛明; 東城幸治
New Entomologist, 64, 13-18, 2015, Refereed, Not invited
Lead

Observation of a Scoprid Stonefly, Scopura montana at the High Altitude Alpine Zone of the Mt. Senjyouga-take and Hijiri-dake, Akaishi Mountains (Japanese Minami ALPS)
東城幸治; 趙在翼; 竹中將起; 三上賢司; 早川美波
New Entomologist, 63, 67-68, 2015, Refereed, Not invited
Lead

The First Observation of the Mating Behavior of the Dipteromimid Mayfly, Dipteromimus tipuliformis (Ephemeroptera, Dipteromimidae)
竹中將起; 東城幸治
New Entomologist, 63, 47-52, 2015, Refereed, Not invited
Lead

Distribution Record of Anisocentropus Caddisfly (Trichoptera, Calamoceratidae) in the Tsushima Islands
Takenaka, M; S. Shibata; T. Suzuki; K. Tojo
New Entomol., 62, 45-49., 2014, Refereed, Not invited
Lead

Winter diet of Japanese macaques from Chubu Sangaku National Park, Japan incorporates freshwater biota
Alexander M. Milner; Susanna A. Wood; Catherine Docherty; Laura Biessy; Masaki Takenaka; Koji Tojo

Development of microsatellite markers for a giant water bug, Appasus japonicus, distributed in East Asia
Tomoya Suzuki; Akira S. Hirao; Masaki Takenaka; Koki Yano; Koji Tojo

Genetic differentiation within species exhibiting widespread gene flow; phylogeography of the downstream-inhabiting speciesEphemera orientalis(Insecta: Ephemeroptera)
Masaki Takenaka; Hiroaki Kogawara; Yeon Jae Bae; Koji Tojo
Lead

Phylogeography of alpine and subalpine adapted Pseudostenophylax caddisflies (Limnephilidae: Trichoptera): a strong relationship with mountain formation
Mikami K; Takenaka M; Nozaki T; Bae YJ; Tojo K

Phylogeography of an insect inhabiting ‘Sky Islands’: the relationships among genetic structures and geographical characteristics, geohistorical characteristics, and cyclical climate changes
Hirohisa Suzuki; Masaki Takenaka; Koji Tojo

高山帯に特異的に生息するヒナバッタにおける遺伝子解析手法の確立および分子系統地理学的研究
竹中 將起; 渡邊 通人; 東城 幸治
昆虫DNA研究会ニュースレター, (30), 26 - 30, Mar. 2019, Not invited
Lead

第2回生物群横断系統地理ワークショップの開催
谷野 宏樹; 竹中 將起
昆虫DNA研究会ニュースレター, (30), 34 - 38, Mar. 2019, Not invited
Lead

The molecular biogeography of aquatic insects in which the influence of geological history is strongly reflected
東城 幸治; 竹中 將起; 谷野 宏樹
生物科学, 70(1), 31 - 41, Jan. 2019, Not invited
Lead

木村資生記念第2回進化学セミナーに参加して
竹中將起
日本進化学会ニュース, 20(1), 24 - 26, 2019, Not invited
Lead

Status update on the metagenomic analysis studies of higher level caddisfly taxa (order Trichoptera)
東城 幸治; 竹中 將起
昆虫と自然, 53(3), 図巻頭1p,18 - 22, Mar. 2018, Not invited
Lead

日本列島に渡来できなかったオオカワゲラ類(カワゲラ目 : オオカワゲラ科Pteronarcyidae)の分子系統地理
竹中 將起; 東城 幸治
昆虫DNA研究会ニュースレター, (26), 13 - 20, Mar. 2017, Not invited
Lead

生物群横断系統地理ワークショップに参加して
竹中將起
日本進化学会ニュース, 18(1), 2017, Not invited
Lead

対馬海峡周辺における昆虫類の遺伝的分断パターンの類型化 : 日本列島・朝鮮半島および対馬の関係性を水生昆虫の遺伝構造から探る
竹中 將起; 柴田 幸紀; 伊藤 富子
昆虫DNA研究会ニュースレター, (23), 42 - 47, Sep. 2015, Not invited
Lead

日本固有科・ガンボカゲロウ科の分子系統地理学的研究
竹中 將起; 東城 幸治
昆虫DNA研究会ニュースレター, (20), 28 - 33, Mar. 2014, Not invited
Lead

水系内に広域分布するチラカゲロウの個体群構造と遺伝的構造-2つの遺伝系統群の検出とそれぞれのハビタット選好性について-
斎藤梨絵; 竹中將起; 東城幸治
日本陸水学会大会講演要旨集, 79th (Web), 2014
Lead

マルヒメツヤドロムシ Zaitzeviaria ovata の分子系統地理
吉田匠, 林成多, 竹中將起, 東城幸治, Joint work
昆虫 DNA 研究会ニュースレター, 2022

日本のオオハサミムシはどこから来たのか?
鈴木智也; 竹中將起; 西川勝; 東城幸治, Single work
昆虫 DNA 研究会ニュースレター, Sep. 2021

全昆虫類に汎用できる DNA バーコーディング領域の探索
竹中將起; 谷野宏樹; 鈴木智也; 東城幸治, Single work
昆虫 DNA 研究会ニュースレター, Sep. 2021

Diversity of sexual traits in insects, unraveled by genes
竹中將起, Editor
昆虫と自然 56, Feb. 2021

連続的な種分化状態を示すガガンボカゲロウを用いた種分化研究
竹中將起; 東城幸治, Single work, 32: 26-31
昆虫DNAニュースレター, 2020

高山帯に特異的に生息するヒナバッタにおける遺伝子解析手法の確立および分子系統地理学的研究
谷野宏樹; 竹中將起, Single work, 30: 34-38
昆虫DNAニュースレター, 2019

第２回生物群横断系統地理ワークショップの開催
谷野宏樹・竹中 將起, Joint work, 30: 34-38
昆虫DNAニュースレター, 2019

地史の影響を強く反映する水生昆虫類の生物系統地理
東城幸治; 竹中將起; 谷野宏樹, Joint work
生物科学, 2019

Status update on the metagenomic analysis studies of higher level Caddisfly taxa (Order Trichoptera)
東城幸治; 竹中將起, Joint work
昆虫と自然, Mar. 2018

The Species and Genetic Diversities of Insect in Japan, With Special Reference to the Aquatic Insects
Koji Tojo; Kazuki Sekiné; Tomoya Suzuki; Rie Saito; Masaki Takenaka, Contributor, Chapter 8 (pp. 229-247)
Springer Japan, 2017

日本列島に渡来できなかったオオカワゲラ類 (カワゲラ目:オオカワゲラ科 Pteronarcyidae)の分子系統地理
Joint work, 26: 13-20
昆虫DNA 研究会ニュースレター, 2015

対馬海峡周辺における昆虫類の遺伝的分断パターンの類型化: 日本列島・朝鮮半島および対馬の関係性を水生昆虫の遺伝構造から探る
竹中 將起 ・柴田 幸紀 ・伊藤 富子・東城 幸治, Joint work, 23: 42-47
昆虫 DNA 研究会ニュースレター 23, 2015

日本固有科・ガガンボカゲロウ科の分子系統地理学的研究
竹中將起; 東城幸治, Joint work, 20: 28-33
昆虫DNA研究会ニュースレター, 2014

Expectations and contributions of phylogeographical study to eDNA study
Masaki Takenaka
環境DNA学会 小集会, Dec. 2023, Invited

冬域の上高地に生息するニホンザルの世界で唯一の採餌行動 −DNA マーカーを用いた越冬戦略の解明−
竹中將起
2023年度信州生態研究会, Nov. 2023, Invited

昆虫類におけるMtInsects-16Sプライマーを用いた環境DNA研究 −従来のCOIプライマーとの比較−
竹中將起
第6回山口大学・環境DNA研究センターシンポジウム, Nov. 2023, Invited

Assessment of genetic diversity in genome-wide studies: phylogeographic and eDNA studies
Masaki Takenaka
神奈川工科大学国際シンポジウム, Aug. 2023, Invited

Behavior of snow monkeys hunting fish to survive winter
Masaki Takenaka
10th Internatinal Charr Symposium (エクスカーション), Jun. 2023, Invited

Environmental DNA study in Insects using the "MtInsects-16S", a newly developed primer for mtDNA 16S region: Achieving highly sensitive detection.
Takenaka M; Hasebe Y; Yano K; Okamoto S; Tojo K
The eDNA Society International Meeting 2023, May 2023, Not invited

厳冬域・上高地におけるニホンザルの生存戦略 -泳ぐ魚のハンティングの観察事例-
竹中將起
2022年度日本動物学会中部支部公開シンポジウム, Nov. 2022, Invited

DNAから紐解く昆虫の不思議な世界 -繁殖生態の進化-
竹中將起; 関根一希; 東城幸治
日本昆虫学会シンポジウム（企画）, Sep. 2022, Not invited

魅力満載！水生昆虫の生物系統地理
竹中將起
第44回日本土壌動物学会 シンポジウム, Jun. 2022, Invited

昆虫の多様な形の進化
竹中將起
昆虫DNA研究会第18回研究集会, May 2022, Invited

種分化連続体を用いた種分化研究 -交配実験における連続的な生殖隔離の検出-
竹中將起
進化学会若手の会, Sep. 2020, Invited

DNAから紐解く昆虫の不思議な世界 -多様性の進化-
竹中將起; 関根一希
Sep. 2019, Not invited

第2回生物群横断系統地理ワークショップ
竹中將起; 谷野宏樹
Jan. 2019, Not invited

集団の分断と生殖隔離-ガガンボカゲロウにおける種分化研究-
竹中將起
種生物学会, Dec. 2018, Invited

DNAから紐解く昆虫の不思議な世界 -昆虫の性の進化-
竹中將起; 関根一希
日本昆虫学会小集会, Sep. 2018, Not invited

EXTREMELY LARGE GENETIC DIFFERENTIATION IN THE MAYFLY DIPTEROMIMUS TIPULIFORMIS: AN IDEAL CASE IN ELUCIDATION OF THE MECHANISMS OF SPECIATION (EPHEMEROPTERA; DIPTEROMIMIDAE)
Masaki Takenaka; Koji Tojo
The 4th Symposium of BSA (Benthological Society of Asia), Aug. 2018, Not invited

Phylogeography of the dipteromimid mayflies which is endemic to the Japanese Islands.
Masaki Takenaka; Koji Tojo
The XV International Conference on Ephemeroptera and XIX International Symposium on Plecoptera, Jun. 2018, Not invited

The reproductive experiment of the Japanese endemic mayfly family Dipteromimidae showing large genetic differentiation: Elucidation of their speciation mechanisms.
Masaki Takenaka; Koji Tojo
The XV International Conference on Ephemeroptera and XIX International Symposium on Plecoptera, Jun. 2018, Not invited

The 3rd Symposium of BSA (Benthological Society of Asia)
Masaki Takenaka; Koji Tojo
Aug. 2016, Not invited

Molecular Phylogeographic Analyses of Anisocentropus Caddisflies (Limnephilidae)
Masaki Takenaka; Saki Shibata; Tomiko Ito; Koji Tojo
15th International Symposium on Trichoptera, Jun. 2015, Not invited

DEEP RELATIONSHIP BETWEEN GENETIC STRUCTURE AND GEOLOGICAL HISTORY OF JAPAN?S ENDEMIC DIPTEROMIMID MAYFLIES, WHICH INHABIT THE UPPER HEADWATERS (EPHEMEROPTERA, DIPTEROMIMIDAE)
Masaki Takenaka; Koji Tojo
Society for Freshwater Science 2015 Annual Meeting, May 2015, Not invited

Molecular phylogenetic analyses of diptemomimid mayflies and its relevance to geohistory: Geological isolation by plate tect onic lines, mountain formation, and Stream Piracies (Insecta, Ephemeroptera, Dipteromimidae)
Masaki Takenaka; Koji Tojo
The 2nd Symposium of BSA (Benthological Society of Asia), Jun. 2014, Not invited

The Phylogeography of Japan’s endemic Dpteromimid mayflies, which inhabit the upper headwaters of rivers (Insecta, Ephemeroptera, Dipteromimidae)
Masaki Takenaka; Koji Tojo
The 2nd Symposium of BSA (Benthological Society of Asia), Jun. 2014, Not invited

Genetic Structures of Dipteromimid Mayflies Endemic to Japan
Masaki Takenaka; Koji Tojo
The First Symposium of BSA (Benthological Society of Asia), Jun. 2012, Not invited

Genetic Structures of Dipteromimid Mayflies Endemic to Japan
Masaki Takenaka; Koji Tojo
International Joint Meeting of the 13th International Conference on Ephemeroptera and 17th International Symposium on Plecoptera, Jun. 2012, Not invited

2020 - Present
日本陸水学会

2019 - Present
昆虫DNA研究会

2014 - Present
THE ZOOLOGICAL SOCIETY OF JAPAN

2013 - Present
THE ENTOMOLOGICAL SOCIETY OF JAPAN

2012 - Present
アジアベントス学会

2016 - 2019
日本生態学会